Wikipedia
Scarabaeinae
The scarab beetle subfamily Scarabaeinae consists of species collectively called true dung beetles. Most of the beetles of this subfamily feed exclusively on dung. Well-known members include the genera Scarabaeus and Sisyphus, and Phanaeus vindex.
Scarabaeinae tribes, with some selected genera and species, are:
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Canthonini
- Anachalcos Hope, 1837
- Circellium Latreille, 1825
- Dicranocara Frolov and Scholtz, 2003
- Versicorpus Deschodt Davis and Scholtz, 2011
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Deltochilum
- Deltochilum valgum
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Coprini
- Copris Geoffroy, 1762
- Heliocopris
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Dichotomiini
- Dichotomius
- Eucraniini
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Eurysterini
- Eurysternus
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Gymnopleurini (sometimes included in Scarabaeini)
- Garreta Janssens, 1940
- Gymnopleurus Illiger, 1803
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Oniticellini
- Euoniticellus Janssens, 1953
- Oniticellus Serville, 1825
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Onitini
- Bubas Mulsant, 1842
- Cheironitis Lansberge, 1875
- Onitis Fabricius, 1798
- Onthophagini
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Phanaeini MacLeay, 1819
- Oxysternon
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Phanaeus
- Phanaeus changdiazi
- Phanaeus vindex
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Scarabaeini
- Pachylomera
- Scarabaeus
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Sisyphini (sometimes included in Scarabaeini)
- Neosisyphus
- Sisyphus
- Unknown tribe / Incertae sedis
- Anisocanthon
- Anomiopus
Coprini, Oniticellini, Onitini, Onthophagini and Phanaeini are sometimes separated as subfamily Coprinae.
scabrosus Fabricius 1776 female Syn. Chironitis scabrosus Fabricius, 1776 (7041225693).jpg| Cheironitis scabrosus indicus Lansberge, 1875 male (2872657334).jpg| Chironitis indicus elphenor Klug 1855 female (4011341553).jpg| Copris elphenor female elphenor Klug, 1855 male (4012106288).jpg| Copris elphenor male