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Alternative clues for the word grouchomarx

• "A Day at the Races" actor
• Antic brother
• Source of the quote
• He said "Here's to our wives and girlfriends ... may they never meet!"
• Speaker of the quip revea
• Speaker of the quip revealed by the ends of the answers to 17-, 25-, 35- and 48-Across
• Comedic brother
• Man with a 36-Across
• I8-#)"'