It can follow the ends of this puzzle's eight longest answers

Usage examples of keepers.

For Leonie, like all Keepers, was a pledged virgin, never to be looked upon with a sensual thought, never to be touched by any man.

Ages of Chaos, when the Keepers were mutilated, even neutered, made less than women.

Oh, yes, the Keepers were still trained, Damon knew, with a terrible discipline, to live apart from men, reflexes deeply built into body and brain.

I know Keepers from the other Towers often leave to marry, but Arilinn?

Neskaya when I was thirteen, and one of the girls told me that if she was sent to Arilinn she would refuse, since the Keepers there were neutered.

Well, that was very long ago, but the tradition remains that these were the first Keepers, neither man nor woman.

You see, most Keepers, if they are going to leave, marry, fall in love, usually leave the Tower before their conditioning is complete.

Then, when love came into your life and you found where your heart lay, then, as has happened with many Keepers, it was no longer possible to remain unawakened.

I was doing the work of three Keepers, and my own health began to suffer.

She had been so proud, that she should succeed where so many failed, that she should be Callista of Arilinn, take her place with the great Keepers of legend.

It was known everywhere in the days when the Towers were as temples, and the Keepers as their priests.

Domains, of those of our family who were Keepers or technicians at Arilinn or Neskaya Tower.

We cannot train enough Keepers as it is now, Damon, and those we train often lose their powers while still young.

Damon lost awareness of his body, it had never existed, he was a dim dream in a dimmer darkness, seeing the faces of Keepers he had never known.

All the Keepers from Nevarsin to Hali are known to me, and there is no Leonie among them, nor no Hastur woman.