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Answer for the clue "Boss on a shield ", 4 letters:
umbo

Alternative clues for the word umbo

Word definitions for umbo in dictionaries

Wikipedia Word definitions in Wikipedia
The umbo (plural umbones or umbos ) is the vaguely defined, often most prominent, highest part of each valve of the shell of a bivalve or univalve mollusk . It usually contains the valve's beak , the oldest point of the valve, and its degree of prominence ...

Wiktionary Word definitions in Wiktionary
n. 1 (context biology English) A boss, or rounded elevation, or a corresponding depression, in a palate, disk, or membrane; as, the umbo in the integument of the larvæ of echinoderms or in the tympanic membrane of the ear. 2 (context zoology English) One ...

WordNet Word definitions in WordNet
n. a slight rounded elevation where the malleus attaches to the eardrum [also: umbones (pl)]

Usage examples of umbo.

The edges were obtuse, the caps fleshy, then corky, smooth, the upper ends not regular, oblique in the form of an umbo or little knob, the pellicles or outside layers thin and easily separated.