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Answer for the clue "Large Carboniferous and Permian amphibians having vertebrae in which some elements remain separate ", 13 letters:
temnospondyli

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Temnospondyli (from Greek τέμνειν ( temnein , "to cut") and σπόνδυλος ( spondylos , "vertebra")) is a diverse subclass of extinct small to giant tetrapods —often considered primitive amphibians —that flourished worldwide during the Carboniferous , Permian ...