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Answer for the clue "A family of fungi closely related to the family Polyporaceae except that the tubes on the undersurface of the cap are separate from each other ", 13 letters:
fistulinaceae

Word definitions for fistulinaceae in dictionaries

Wikipedia Word definitions in Wikipedia
The Fistulinaceae are a family of fungi , the best-known member of which is the beefsteak fungus Fistulina hepatica . Molecular studies have now shown it to lie within the Agaricales .