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Answer for the clue "A large and taxonomically difficult group of Ascomycetes in which the fleshy fruiting body is disklike or cup-shaped ", 12 letters:
discomycetes

Word definitions for discomycetes in dictionaries

Wikipedia Word definitions in Wikipedia
Scarlet Elf cup. Discomycetes is a former taxonomic class of Ascomycete fungi which contains all of the cup, sponge, brain, and some club-like fungi. It includes typical cup fungi like the scarlet elf cup and the orange peel fungus , and fungi with fruiting ...