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Answer for the clue "A class of mostly aquatic fungi ", 16 letters:
chytridiomycetes

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Chytridiomycetes is a class of fungi . Members are found in soil, fresh water, and saline estuaries. They are primitive fungi, and are first known from the Rhynie chert . It has recently been redefined to exclude the taxa Neocallimastigomycota and Monoblepharidomycetes ...