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Answer for the clue "An orange of low brightness and saturation ", 9 letters:
brownness

Word definitions for brownness in dictionaries

The Collaborative International Dictionary Word definitions in The Collaborative International Dictionary
Brownness \Brown"ness\, n. The quality or state of being brown. Now like I brown (O lovely brown thy hair); Only in brownness beauty dwelleth there. --Drayton.

WordNet Word definitions in WordNet
n. an orange of low brightness and saturation [syn: brown ]

Wiktionary Word definitions in Wiktionary
n. The characteristic of being brown.

Usage examples of brownness.

Beneath his window was a neat garden, scruffy now with autumn brownness, in the center of which was a formal quatrefoil pond, the water green with algae and rippling in the brisk wind.

To illustrate with the example you've offered -- brown hair versus blond -- let's imagine a dominant gene, B, for brownness, and its recessive counterpart, W, for whiteness.