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Answer for the clue "An orange of low brightness and saturation ", 9 letters:
brownness

Word definitions for brownness in dictionaries

Wiktionary Word definitions in Wiktionary
n. The characteristic of being brown.

WordNet Word definitions in WordNet
n. an orange of low brightness and saturation [syn: brown ]

The Collaborative International Dictionary Word definitions in The Collaborative International Dictionary
Brownness \Brown"ness\, n. The quality or state of being brown. Now like I brown (O lovely brown thy hair); Only in brownness beauty dwelleth there. --Drayton.

Usage examples of brownness.

Beneath his window was a neat garden, scruffy now with autumn brownness, in the center of which was a formal quatrefoil pond, the water green with algae and rippling in the brisk wind.

To illustrate with the example you've offered -- brown hair versus blond -- let's imagine a dominant gene, B, for brownness, and its recessive counterpart, W, for whiteness.