Wikipedia
The extinct five-horned prongdeer Hoplitomeryx matthei with its sabrelike (' moschid' type) upper canines lived on the former Gargano Island during the Miocene and the Early Pliocene, now a peninsula on the east coast of South Italy.
Its fossilized remains were retrieved in the late sixties and subsequent years from reworked reddish, massive or crudely stratified silty-sandy clays (terrae rossae), which partially fill the paleo-karstic fissures in the Mesozoic limestone substrate and that are on their turn overlain by Late-Pliocene-Early Pleistocene sediments of a subsequently marine, shallow water and terrigenous origin. In this way a buried paleokarst originated.
The fauna from the paleokarst fillings is known as Mikrotia fauna after the endemic murid of the region (initially named "Microtia", with a c, but later corrected, because the genus Microtia was already occupied). Later, after the regression and continentalization of the area, a second karstic cycle started in the late Early Pleistocene, the neokarst, which removed part of the paleokarst fill.