Wiktionary
cosmine
n. A continuous layer of dentine, covered by a layer of enamel, that takes the place of scales in some ancient fish.
Wikipedia
Cosmine
Cosmine is a spongy, bony material that makes up the dentine-like layers in the scales of the lobe-finned fishes of the class Sarcopterygii. Fish scales that include layers of cosmine are known as cosmoid scales.